glossary iconglossary

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glossary icon - A

Abortion Halt in development of an organ after partial differentiation.
Abscission Premature drop off or falling of leaves, flowers, or fruit resulting from abnormal development of anthocyanin pigments.
Acanthae Cuticular projections on the surface and in the foregut of insects
Accessory doublet Ring of small microtubules surrounding the central tubules in the axoneme of the sperm
Aeciospore A spore borne in an aecium; dikaryotic; separated by disjunctor pads
Aecium Cup-shaped sorus of the Uredinales producing aeciospores (pl. aecia)
Acephalic Head capsule almost completely retracted into the thorax
Acervuli Saucer-shaped structure with conidia on short conidiophores
Acetabulum Cavity in the integument into which the protuberance of an appendage is fitted
Acid gland Gland which produces venom in Hymenoptera
Acrosome An organelle found at the apex of the head of the sperm
Acrosome reaction Leakage of enzymes from the sperm which enables egg penetration
Acrotergite Narrow sclerite on the anterior margin of the notum
Acrotrophic ovary See telotrophic ovary
Active ingredient The ingredient in a pesticide formulation that has the pesticidal effect.
Adecticious Pupal form with non-functional mandibles
Adephaga Suborder of the Coleoptera (beetles) e.g. ground beetles, tiger beetles
Aedeagus Distal part of the phallus, principal part of the intromittent organ
Aeropyle Structure of the eggshell responsible for respiration
Agar A substance produced from certain red algae used to make culture media into gels
Agronomy The theory and practice of crop production and soil management.
Alinotum Anterior wing-bearing region of the pterothoracic notum
Allelopathic A chemical process that a plant uses to keep other plants from growing too close to it
Allocryptic Resembling familiar objects in its environment for concealment
Allopatric speciation Allo = Greek for 'other', 'another'; patria = Greek for 'country' (fatherland). Speciation between geographically separated populations
Allospecific From or of another species
Allosematic Resembling the colouration of dangerous or inedible animals or plants for protection
Alternative host A host other than the normal (crop) host on which the pathogen can exist
Alien plant A plant from another country
Ametaboly Lacking metamorphosis; adult resembles the immature stages
Amphigynous Where the antheridium surrounds the oogonial stalk (in certain Pythiaceae)
Amplexiform Wing coupling where the jugum of the forewing overlaps the humeral lobe of the hindwing
Anagenesis Evolution of a feature or features within a lineage, including the gradual evolution into a ‘new’ species without splitting into two taxa
Anagenesis Process of progressive evolution
Analogy Structures in two different organisms with the same function
Anal veins (vannal veins) Associated with the third axillary sclerite
Anamorphosis Development of additional abdominal segments in the adult stage
Androconia Specialised scale in Lepidoptera that disseminate scents
Angiosperms Flowering plants
Anisoptera Suborder of Odonata (dragonflies)
Annuals A plant which grows, produces seeds, and dies within one year
Anocellate Lacking ocelli
Antecosta Apodemal projection
Antecostal suture Suture separating the acrotergite and alinotum
Antennal mechanosensory and motor centre Found in the middle region of the brain and innervates the antennae where it is responsible for mechanoreception
Anterior notal wing process Anterior articulation point for the wing
Anthesis The period of flowering
Anthocyanescence A reddish or purplish colouration of leaves resulting from abnormal development of anthocyanin pigments.
Anthophilous Feeding on flower parts
Anthracnose Dark, sunken, necrotic spots or patches
Apical (=distal) Structure or part of a structure furthest from the body
Apneustic respiratory system Respiration is through the integument and spiracles are closed
Apodeme Cuticular inflection of the body wall marked externally by pits
Apocrita Suborder of Hymenoptera (bees, wasps, ants) e.g. ants, wasps
Apodous Larval type without legs
Apolysis Stage of moulting where the old and new cuticle separate
Apomorphic Derived or recent condition
Apophyses Arm-like apodemes
Aposematic Colouration warning of unpalatibility
Apothecium Saucer or cup-shaped ascocarp (pl. apothecia)
Appendix dorsalis Long median filament at the end of the abdomen
Apposition eye Forms an image when light reaching the rhabdom enters through the lens of that same ommatidium. Typical of diurnal insects
Appressoria
(sing. appressorium)
Swellings on germ tubes or hyphae, rounded and either simple or lobed, adhere firmly to surfaces by their mucilaginous walls. Formed as holdfasts from which fine infecting hyphae can be forced down through protective structures such as the cuticle overlying epidermal cells.
Apterous Insect which is totally wingless
Aptery Condition where insect is totally wingless
Apterygotes Hexapod arthropods that are primitively without wings, now usually considered an artificial group (paraphyletic) - Archaeognatha and Thysanura
Arolium A lobe of the pretarus arising between the claws.
Archeognatha Order of insects in the Subclass Apterygota: bristletails
Arrhenotokous Producing male offspsring from unfertilised eggs
Ascocarp An ascus-producing structure in the Ascomycota
Ascomycota The group of fungi which produce ascospores in asci and is the largest fungal phylum
Ascospores Spores produced by the Ascomycota which are borne in asci
Ascus Typical sexual reproductive cell of the Ascomycota; usually contains eight haploid ascospores (pl. asci)
Aseptate hypha Hyphae which do not have septa
Asexual spores Spores that are as a result of vegetative reproduction (similar to budding)
Assemblage A group or collection of taxonomically related species living in a particular area
Assembly zone Above the epidermal cells and beneath the cuticle in the integument
Attenuate Becoming progressively thinner
Auchenorrhyncha Suborder of the Hemiptera (bugs) e.g. cicadas, spittlebugs, leafhoppers, treehoppers
Autapomorphy A derived character unique to a single taxon, useful for diagnosing a taxon, but lacking information on related taxa. For example, all scorpions can be recognised by their unique terminal sting, but this character provides no information about their relationship to other chelicerates.<a name="b" id="b"></a>
Autotomy Shedding of appendages for defence
Autotroph ‘self nourishing’ — organisms (e.g. Green plants) that produce their own food from inanimate nature; opposed to heterotroph
Axillary region Basal area of the wing; contains the axillary sclerites.
Axoneme Functional unit of the flagellum consisting of microtubules
Axon Principle nerve fibre of a neurone

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glossary icon - B

Bark roughening Formation of rough bark where smooth bark normally occurs.
Basalare A sclerite at the base of the wing anterior of the pleural wing process.
Basal pigment cells When present, surrounds the base of the retinular cells
Basement membrane Bottom layer of the integument
Basidiomycota Phylum of fungi characterised by the formation of club-shaped structures termed basidia which each produce four haploid basidiospores. Includes fungi most familiar to people eg. mushrooms
Basidiospore A (haploid) spore produced by a basidium.
Basidium Typical sexual reproductive cell of the Basidiomycota; usually produces four haploid basidiospores (pl. basidia)
Basisternum Anterior sclerite of the mediosternite.
Batesian mimicry Harmless edible species resemble harmful, inedible species
Bauplan Body plan, the basic structural organisation of a group of organisms
Biocontrol Introduction of natural enemies (parasites, predators, and/or pathogenic microorganisms) to reduce or suppress populations of plant or animal pests
Biodiversity The diversity of life, including genes, populations, species, assemblages and communities
Biomass A renewable energy resource derived from the carbonaceous waste
Biotrophic Living in balance with the host, at least initially
Biotrophy Gain nutrients from living host cells by means of haustoria
Biramous Limbs with ‘two branches’, as in the primitive arthropod limb
Blackleg Necrosis of stem tissues causing a darkening and rotting of stem bases
Blasting Failure to develop fruit.
Blattodea Order of insects: cockroaches
Bleeding Sudden drying and browning of whole leaves, shoots or branches.
Blight Sudden and extensive shrivelling and death of certain areas of the plant; coalescence of many individual lesions
Brachypterous Condition where wings are disproportionately small in relation to the body
Brachyptery Condition where wings are disproportionately small in relation to the body
Brachycera Suborder of Diptera (flies) e.g. march flies, hover flies, houseflies, blow flies.
Branchial chamber Enlarged anterior part of the rectum forms a chamber which contains a system of tracheal gills, the branchial basket. Found in dragonflies.
Buccal cavity Region of the foregut immediately inside the mouth
Burn A synonym for scorch.
Bursa copulatrix A sac-like vagina

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glossary icon - C

Caeca A blind sac or pouch attached to the gut
Caelifera A suborder of Orthoptera (grasshoppers, crickets) e.g. short horned grasshoppers
Calcar A spur used for grooming in Hymenoptera
Callus Overgrowths of tissue produced in response to injury or other irritation and which tend to cover a wound, canker etc.
Calyx Enlargement of the lateral oviduct near the pedicel of the ovariole
Canker A sunken necrotic lesion beginning in the bark of branches, trunk or roots and usually becoming deeper and wider by normal or accelerated secondary growth (callusing) of cambium, phellogen, and traumatic meristem.
Campodeiform Larva which is elongated, flattened with long well-developed legs
Capitate Having a knob or head at the apex.
Capitulum Cap-like structure on the operculum
Cardo Basal segment of maxilla; bears stipes
Carinae Elevated cuticular ridges
Carotenoids Pigments which produce yellow and orange colours
Caudal gill A type of tracheal gill which occurs in some Odonata
Caudal visceral nervous system Nervous system that innervates the posterior part of the hindgut and reproductive organs
Caudal style See appendiix dorsalis
Central nervous system Central series of ganglia extending the length of the body
Central tubules Conspicuous elements of the axoneme
Cerci Abdominal appendages which originate on segment 11
Chelicerae The most anterior feeding appendages in the Chelicerata, usually pincer-like, but may be modified into fangs (e.g. Spiders) or stylets (e.g. Spider mites)
Chlamydospore A thick-walled asexual spore formed by the rounding up of an existing cell
Chlorophyll The green pigment in plants that absorbs energy from sunlight necessary for photosynthesis.
Chlorosis Yellowing of leaves as a result of inhibition of chlorophyll production. Often caused by nutrient deficiencies
Chlorotic spots Yellow spots on leaves as a result of inhibition of chlorophyll production. Often caused by nutrient deficiencies
Chordotonal organ Subcuticular sound receptor
Cibarial pump A pumping mechanism adapted for imbibing fluids e,g, Homoptera
Cibarium Dorsal food pouch in the preoral cavity (formed by the hypopharynx)
Characters Anatomical features
Character states Stages of expression of the characters
Chitin Major component of cuticle
Clades Branches on the tree on life; monophyletic groups
Cladistics Phylogenetic systematics, the discovery and use of monophyletic groups
Cladogenesis The splitting pattern produced by speciation; the generation of clades
Cladogram A picture (diagram) of phylogenetic relationships; a representation of a hypothesis about the relationships among monophyletic groups
Clavate Club-shaped
Cleistothecium An ascocarp with an enclosed globuse structure with no natural opening that ruptures when mature
Clypeus Sclerite immediately above labrum
Coarctate Pupal development occurs in a puparium
Coeloconic sensillum Sense organ sunk into a pit or depression in the integument
Coenocytic Possessing a protoplast which is multinucleate and/or without cross walls
Coleoptera Order of insects: beetles. Greek coleos = sheath; pteron = wing
Collaterial gland Gland that secretes adhesive substances
Collembola Class of entognathous hexapods: springtails. Greek, kolla = glue; embolon = peg.
Collophore Appendage on the 1st abdominal segment in Collembola
Community All interacting species that occur in a given area; it is very rare to study a complete community, but common to study assemblages
Competition An interacation between plants that consume resources present in a limited supply
Competitive cultivars Cultivars which compete with weeds
Competitive crop species Crop species which compete with weeds, suppressing weed growth and weed seed production while maintaining acceptable grain yields.
Complementarity A way of viewing two species assemblages that emphasises the differences between them; the ratio of the number of species unique to a collection to the total number of species in two collections
Condyle A knob-like process which articulates the mandible to the head
Conidiophore Asexual spore exposed directly to the atmosphere and borne on conidiophores (pl. conidia)
Conidium simple or branched hyphae
Conspecific Of the same species
Containment Keep the harmful weeds within limits and reduce the rate of spread of a new weed
Control decision Pest control decision made that relies on a cost to benefit assessment of the pest situation
Convergent evolution Structures resembling one another without a common origin
Coprophagy Eating faeces
Coralloid Having the form or consistency of coral
Corbiculae Pollen basket found on the coxa of Hymenoptera
Cornicles Secretory structures on the abdomen of aphids
Costal vein Dominant vein near or confluent with the anterior margin of the wing
Coxa First segment of the leg, attached to the thorax
Coxopodite Base of the primitive insect leg now represented by the coxa
Crepuscular Indicates an organism is active at dusk or dawn
Crystalline cone Immediately beneath the lens; composed of a translucent material.
Crop Part of the foregut where food is stored
Crop rotation Crops are changed year by year in a planned sequence
Cropping system The system used to grow and manage crops for particular end-usesIs
Crop topping Applying herbicides late in the season to control weed seed set
Crotchets Curved hooks or spines on the prolegs of Lepidoptera
Cryptic An insect imitates a plant part or inanimate object
Crypsis Insect imitating a plant part or inanimate object
Cryptogams ‘hidden reproducers’, mosses, liverworts and lichens
Ctenidia Rearward projecting cuticular spines on fleas
Cubital vein Fifth vein, associated with the distal median plate
Curl (roll) Abnormal bending or curling of leaves or shoots due to localized overgrowth on one side or in certain tissues.
Cursorial Adapted for running
Cuticle Outer most layer of the integument
Cuticular intima Inner most layer of the gut
Cuticulin layer Layer of the epicuticle; barrier between the old and new cuticle
Cyclopean ear Hearing organ in mantids; found on the metathoracic leg
Cytochrome A family of over 60 enzymes the body uses to metabolise toxic hydrocarbons, and make blood.

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glossary icon - D

Damping-off A symptom complex characterised by rapid dying, browning, and rotting of germinating seedlings.
Decticious Pupal form with articulated, functional mandibles
Defoliation Loss of leaves through abscission.
Dendrite Branches given off from a nerve cell
Dens Elongate conical structures of the furcula
Dermal glands Modified epithelial cells found in the epidermis
Dermaptera Order of insects: earwigs. Greek, derma = skin, pteron = wing.
Detritivore Detritus eating
Deutocerebrum Middle section of the brain
Deuteromycota Phylum of fungi which reproduce only by asexual means
Diapause Anticipated delayed development which occurs annually
Dichoptic Eyes not large and do not meet medially
Dicondylic mandible Mandible with two articulation points with the head
Dieback Progressive drying, shrivelling, and browning of twigs or branches from the tips inward toward the trunk.
Digitate Finger-like
Diplura Class of entognathous hexapods. Greek, diploos = tail; oura = tail
Diptera Order of insects: flies. Greek, dis = twice; pteron = wing
Distal median plate At the base of the mediocubital region, often absent
Ditrysian Female Lepidoptera with 2 abdominal apertures associated with the reproductive system
Drop Premature abscission of fruit or twigs.
Dropsy Extensive intumescence over large areas of tissue.
Dufours’s gland Accessory gland found in Hymenoptera
Dwarfing (nanism) Subnormal size of a plant or some of its organs.

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glossary icon - E

Ecdysial suture Longitudinal suture on head; important in moulting
Ecdysis Final stage of moulting where the old cuticle is shed
Eclosion The emergence of immature stage from the egg i.e. hatching
Economic loss Financial loss to a grower resulting from lower yield or reduced quality of a grown crop
Economic threshold The pest density at which management intervention must be taken to prevent the pest from reaching the economic injury level.
Ecosystem A naturally occuring assemblage of organisms (plant, animal and other living organisms) living together with their environment, functioning as a unit of sorts.
Ectognatha Hexapods whose mandibles remain exposed and are not extensible. These are the true insects.
Ejaculatory duct Duct formed by the joining of the vas deferens from each testis
Elytra Modified, hardened forewing in beetles that protects the hind wing
Embioptera Order of insects: webspinner.
Empodium Median lobe or spine arising ventrally between the pretarsal claws
Endocuticle Inner most layer of the cuticle
Endophallus Inner chamber of the phallus invaginated at the end of the aedeagus into which the ejaculatory duct opens
Endopterygota Insects which develop wings internally
End-twigging End branching of wing veins in dragonflies (Odonata)
Ensifera Suborder of Orthoptera (grasshoppers & crickets) which are the long-horned grasshoppers e.g. king crickets, katydids, wood crickets, mole crickets.
Entognatha Hexapods which maintain their andibles concealed when not in use e.g. Protura, Collembola and Diplura
Entomophagous Insect-eating
Environmental weeds Plants that invade native communities or ecosystems
Ephemeroptera Order of insects: mayflies. Greek, ephemeros = lasting a day; pteron = wing
Epidermis Layer immediately above the basement membrane
Epididymis Highly convoluted seminal vesicle
Epimeron Posterior sclerite of the pleuron.
Epimorphosis Fixed number of body segments in the embryo before hatching
Epinasty A leaf curl in which the leaves turn downward, due to a more rapid growth of cells on the upper side than on the lower side of the leaf stalk.
Epiproct Dorsal surface of the 11th abdominal segment
Episternum Anterior sclerite of the pleuron.
Epitoky Successive alternation of a feeding, non-reproductive instar with a reproductive instar during the adult stage
Eradication To get rid of completely
Eruciform Larval type which is caterpillar-like, cylindrical with short legs
Etiolation A symptom complex in which the major symptoms are dwarfing of foliage and inflorescence, spindly stem growth, and chlorosis.
Etiology The study of the cause of disease
Eucephalic Fully exposed, well-developed head capsule
Eukaryote Having intracellular, membrane-bound organelles
Eusocial Insects that display social characteristics e.g. Termites
Eusternum Anterior region of the sternum - divided into presternum, basisternum and sternellum.
Exarate Pupal form with free appendages
Exite theory The theory that wings evolved from pre-existing coxal exites
Exocuticle Outer layer of the cuticle
Exopterygota Insects whose wings develop externally

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glossary icon - F

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glossary icon - G

Fasciation Flattening or cohering of organs such as stems, flowers, and roots. The cause is unknown, but in some cases the condition may be propagated.
Fasciculation (brooming) The clustering of organs such as branches about some common point on the plant. Generally arises from development of adventitious buds.
Fecundity Number of eggs produced by a female
Femur Third and largest segment of the leg ("thigh")
Fertility Number of viable eggs
Filter chamber Found in many plant-fluid feeding bugs; an adaptation of the midgut enabling the insect to absorb essential nutrients from the large quantity of fluid absorbed
Fimbriae A structure that can aid the attachment of bacterial cells to plant surfaces; believed to be a means for genetic exchange between bacterial cells by conjugation
First axillary sclerite Hinge plate of wing base, associated with the subcostal
Flagellum Tail of the sperm
Flagging A permanent condition of wilting.
Follicle Funcitonal unit of the testis
Follicle cells Inner epithelial cells of the ovariole
Fossorial Adapted for digging
Frass Solid larval excrement
Frenate Coupling Bristles of the hindwing hook under the base of the forewing
Frons Part of head immediately ventral to vertex
Frontoclypeal suture Transverse suture beneath the antennae; separates frons and clypeus
Fungivorous Feeding on fungus
Furcula Jumping organ of Collembola on 4th abdominal segment
Furcal pits Paired external pits on the sternacostal suture

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glossary icon - G

Galea Outer lobe of the maxilla; attached to the stipes
Gall A pronounced tumefaction, often more or less spherical and usually composed of undifferentiated cells.
Gamete A reproductive cell that is capable of fusing with another reproductive cell
Ganglion A mass formed by groups of nerve cells
Gaster Remaining segments of abdomen in Hymenoptera
Gena "Cheek" of insect; on side of the head below the eyes
Gene flow Migration of genes from one population to another, through movement and successful reproduction
Gene frequency A measure of the occurrence of an allele in a population.
Germarium Apical region of the testis or ovariole, responsible for producing the germ cells
Germ pore A thin area in a spore wall through which a germ tube may emerge
Girdling Tangential enlargement of a canker or lateral coalescence of cankers causing a branch or stem to be encircled in the complete stoppage of conduction.
Glial cells Cells which surround the neurone and isolate the axon
Glossae Inner lobes on prementum (part of labium)
Glycocalyx Outer covering of the mature spermatazoon
Goblet cells Interspersed between the epithelium cells of the midgut; function in maintaining the water balance
Gonapophysis Genital stylus on the abdomen of Thysanura
Gonopore Female - aperture through which the egg passes during oviposition
Male - opening of ejaculatory duct into the endophallus
Gonostyli Accesory clasping structure
Graminicides Herbicides for controlling grasses
Gram stain A stain of bacteria yielding either a positive or negative reaction depending on the amino acid composition of the bacterial cell wall
Gressorial Adapted for walking
Groundplan Hypothetical primitive condition
Guilds A group of species living in the same place that is using the same habitat in a similar way
Gula Structure formed by fusion of the postocciput
Gummosis Formation of gums by diseased cells and tissues and the extrusion of gum from wounds and other lesions.
Gymnosperms ‘naked seeds’, the conifers and their allies

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glossary icon - H

Haemocoel The large body cavity in which haemolymph flows
Haemolymph Insect "blood"
Hair plates Concentrations of setae found on many parts of the body
Hairy root A type of fasciculation affecting the roots.
Haltere Modified hindwing of Diptera; used for balancing
Hamulate coupling Hooks (hamuli) on the costal region of hindwings engage with the anal region of forewings
Harpagones Accesory clasping structures
Hastisetae Setae which are used in defense in larval carpet/hide beetles
Haustellate Having mouthparts adapted for sucking e.g. bugs
Haustorium A hyphal branch which penetrates a host cell and absorbs nutrients from it (pl. haustoria)
Hemelytron Forewings in Heteroptera, with a hardened basal region
Hemi-biotrophy Gain nutrients by both biotrophic and necrotrophic means
Hemicephalic Head capsule more or less reduced or incomplete posteriorly
Hemimetabolous Life cycle where there is one conspicuous radical change in form from immature to adult
Hemiptera Order of insects: bugs. Greek, hemi=half; pteron = wing
Herbicide Chemicals that kill plants principally by interfering with the normal biochemical processes of plants
Herbicide resistance The inherited ability of a plant to survive and reproduce following exposure to a dose of herbicide that would normally be lethal to the wild type
Herbivore An animal that gets its energy from eating plants, and only plants
Heterogony ‘Cyclical production of males
Heteroptera ‘Suborder of the Hemiptera (bugs) e.g. stink bugs, assassin bugs, jewel bugs
Heterotrophy Adventitious development of normal tissues or organs in unusual areas.
Heterotroph ‘different nourishing’ – organisms (e.g. Animals) that obtain food from other organisms; opposed to autotroph
Histogenesis Development and formation of tissue
Histolysis Breakdown and dissolving of tissue or internal structure by the haemolymph
Holometabolous Life cycle where there is a pupal stage between immature and adult stages
Holoptic Large eyes which meet medially
Homology The study of the same structure or gene in different organisms
Homoplasy Phylogenetic noise; mistaken homologies due to convergence
Honeydew A watery fluid containing sugars eliminated from some Homoptera
Humeral plate Anterior sclerite at the wing base, supports the costa
Humeral vein Cross vein connecting the costa and subcosta.
Hyaline Clear, colourless
Hydathode A structure (usually in a leaf) that permits the release of liquid water through a pore in the epidermis
Hydropyle Device which absorbs water
Hydrosis A water-soaked, translucent appearance of leaves, fruits and green stems due to the extrusion of water from the cells into the intercellular spaces.
Hymenium Sporing layer of a fruiting body (pl. hymenia)
Hymenoptera Order of insects: ants, bees and wasps. Greek, hymen = membrane; pteron = wing
Hypandrium Ninth segment of males in some insects
Hypergamesis Process where excess sperm are used by the female
Hypermetamorphosis Undergoing major morphological changes between larval instars
Hyperparasitoid Parasitoid of a parasitoid
Hyperplasia An abnormal increase in the number of cells in a particular tissue or organ.
Hypertrophy An abnormal increase in the size of a plant or plant part due generally to an abnormal increase in the size of the cells.
Hypha Multicellular branching filaments of fungi (pl. hyphae)
Hypocotyl The part of a seedling below the cotyledons and above the root
Hypognathous Mouthparts oriented vertically
Hypopharynx Lobe which extends from back of preoral cavity; divides preoral cavity horizontally
Hypostomal bridge Fusion of the posterior region of the subgena

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glossary icon - I

Incursion The appearance of weeds in an area where they had not occurred before
Infestation Presence of a weed (for example) in large numbers thus causing problems
Infection court The place on or in the host where infection takes place
Inflection Cuticular ridges with a precipitous elevation
Instar Immature insect between moults
Integrated crop management An holistic approach to planning and execution of profitable crop production, taking into consideration climatic and soil factors, crop rotations, crop variety, wildlife conservation, energy conservation, livestock enterprises and Integrated Pest Management.
Integrated weed management The use of a combination of different agronomic practices to manage weeds, so that the reliance on any one weed control technique is reduced.
Intumescence Small epidermal swellings or eruptions resulting from hypertrophy of groups of epidermal or sub-epidermal cells due to high turgor pressure.
In vitro Literally "in glass" i.e. in culture
Inundative biocontrol Regulation of populations of a target weed by the repeated introduction of natural enemies (often fungi) of the weed
Invasion When a weed enters an area and then dominates the native species
Invasive weed A weed that has rapidly moved into an area and is difficult to stop and control
In vivo In a live host
Isoptera Order of insects: termites. Greek, iso = equal; pteron = wing
Integument Multi-layered outer covering of an insect
Interneurones Unipolar neurones that lie in the CNS and direct nerve impluses

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glossary icon - J

Johnston’s organ Auditory organ located within the 2nd antennal segment (pedicel)
Jugal veins In the jugal region; responsible for flexing the wing
Jugum Membranous lobe, proximal of the anal region,
Jugate Coupling Wing coupling mechanism - jugum of the forewing covers the costal margin of the hindwing

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glossary icon - K

Knots A type of tumefaction.

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glossary icon - L

Labella Fleshy, lobe-like structure formed by labial palps in flies
Labium Lower lip of the mandibulate mouth
Labrum "Upper lip" of insect; beneath clypeus
Lacinia Inner lobe of maxilla; attached to stipes
Lamellate gills Leaf-like gills
Larva Immature insects of holometabolous insects
Laterosternite The separated lateral part of the sternum found in some insects
Latrotergites Sclerites of the lateral portion of the dorsal surface
Leaf retention Abnormally long retention of leaves usually resulting from a failure to develop the abscission meristem.
Legislation A law or set of laws suggested by a government and made official by parliament for the regulation of weed species
Lens Transparent, usually colourless, dome-shaped cone cells; focus light
Lenticel A region in the phellem characterised by intercellular spaces
Lepidoptera Order of insects: butterflies and moths. Greek, lepido = scale; pteron = wing
Lesion A localized area of diseased tissue.

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glossary icon - M

Macrochaetae Long, non-articulated setae on the forewing of Trichoptera
Macroptery Condition where wings are fully developed
Macrochaetae (=macrotrichia) long, non-articulated setae on the forewing of Trichoptera
Macrophytes Large plants; opposed to microphytes, e.g. Single-celled algae
Malpighian tubules Organs of excretion found at the junction of mid- and hindgut.
Mandible Paired sclerotised "jaws"
Mantodea Order of insects: preying mantids
Manubrium Medial component of the furcula
Maxilla Paired, segmented secondary jaws; between mandibles and labium
Meconium Substance excreted by prepupa or pupa or by newly emerged adults of certain insects
Mecoptera Order of insects: scorpion flies. Greek, mecos = long; pteron = wing
Medial cross vein Connects secondary branches of median, M2 and M3.
Medial vein Fourth vein; associated with the median plates
Medio-cubital cross vein Connects posterior most branch of median with anterior most branch of cubital
Megaloptera Order of insects: alderflies. Greek, megas = large, pteron = wing.
Melanins Pigments which produce brown and black colours
Meron A posterior subdivision of the coxa which dissociates from the coxa and becomes incorporated into the pleural wall.
Mesocuticle Transitional cuticular region associated with the endocuticle
Mesosoma Combined propodeum and thorax in Hymenoptera
Mesospore A 1-celled teliospore among 2-celled ones
Mesothorax Second segment of the thorax
Metamerism The organisation of an arthropods body as a serially repeated series of body segments
Metamorphosis Changes in body form from immature to adult
Metasoma Remaining abdominal segments in Hymenoptera
Metathorax Third segment of the thorax
Micropyle Minute aperture in the chorion of the egg
Microchaetae (=microtrichae) Short, non-articulated setae on forewing of Diptera; hair-like structures similar to setae but are fixed
Microchaetae One of a number of small tubular structures found inside nearly all cells used for the transport of materials inside cells
Microtubules A hollow cylindrical tube found inside nearly all cells and used for the transport of materials inside cells.
Mitosis The process of nuclear division in cells that produces daughter cells that are genetically identical to each other and to the parent.
Monocondylic mandible Mandible with a single articulation point with the head
Monophyletic A group of taxa all descended from the same common ancestor, and including all descendents of that ancestor; a clade
Monotrysia Female Lepidoptera with 1 terminal abdominal aperture
Morphogenesis Anatomical change of structure during development
Morphospecies ‘form species’ an OUT based on general appearance, without confirmation of species identity. Also, a species diagnosed on strictly morphological characters.
Mosaic Pale green mottling of leaves.
Motor neurone Innervate and stimulate muscles
Moult Process of new cuticle formation and the shedding of the old cuticle
Mucron Sharp pointed structures of the furcula
Muellerian mimicry Two or more distasteful or harmful species mimic each other
Mulch A protective layer of a organic or inorganic material that is spread on top of the soil e.g. pine bark, straw, plastic, stones
Mulching To place a protective layer of a material on top of the soil to protect the soil, reduce water loss etc.
Multivoltine Insects with many generations in a year or season
Mummy A dried shrivelled fruit that is generally the result of a fungus disease.
Mycophagous Fungus eating
Mycelium A mass of hyphae (pl. mycelia)
Mycorrhiza Fungal symbiotes associated with plant roots

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glossary icon - N

Natatorial Adapted for swimming
Native pasture Pasture made up predominantly of native grasses
Native plants Plants which grow naturally in a place, and have not been brought there from somewhere else
Necrophagous Feeding on dead or decaying matter
Necrotrophic Only obtain nutrients from dead host cells, they kill host tissue
Necrosis Death and disintegration of cells and tissues.
Needle cast Abscission or abnormal loss of needles from those species which retain them.
Nematocera Suborder of Diptera (flies) e.g. crane flies, mosquitoes
Nematode An unsegmented worm, belonging to the phylum Nematoda, which is generally round and pointed at both ends. There are both parasitic and free-living species.
Neoptera Insects with wings folded over the abdomen when at rest, occurs in most insect orders.
Neotenics Secondary reproductives of the Isoptera (termites)
Neurone Nerve cell including axon and dendrites. It is the basic functional unit of the nervous system
Neuroptera Order of insects: lacewings. Greek, neuro = nerve; pteron = wing
Nidi Groups of regenerative cells in the midgut
Nitrophilous A plant growing best in a habitat rich in nitrogen.
Nodal points Crticial points in the history of insects which reflect evolutionary change
No-tillage Planting crops without prior seedbed preparation, into an existing cover crop, sod, or crop residues, and eliminating subsequent tillage operations.
No-till cropping No-till cropping is a system where the aim is to avoid tillage
Notum Dorsal sclerite of thorax (= tergum of abdomen)
Nudisetae Smooth setae distributed over the body
Nymph Immature insect which emerges in a relatively advanced stage of development

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glossary icon - O

Obtect Pupal form with appendages "glued" to the body
Ocelli Simple eyes found on the vertex in most adult insects. Enhances light detection, changes in light intensity
Occipital foramen Opening at back of head through which internal structures enter thorax
Occiput Back of the head between the vertex and neck
Odonata Order of insects: dragonflies. Greek, odous = tooth, pteron = wing
Oligomerisation Division of functional responsibility
Oligoneustic respiratory system Respiratory system characterised by 1 or 2 pairs of spiracles. Two types of oligopneustic systems are recognised: amphipneustic and metapneustic
Oligopodous Larval type with 3 thoracic legs but no prolegs present
Ommatidium Basic unit of vision; a compound eye is made up of a number of ommatidia
Ommochromes Pigments which produce yellow, red and brown
Ontogeny Pathways of embryonic development
Oo (pref.) - egg
Oocytes Egg cells formed by the oogonia
Oogonia First stage germ cells in the female which produce the oocytes
Oogenesis Development of the oocyte
Oomycota A phylum which are not true fungi but rather are members of the plant kingdom Chromista
Oospores Sexual spores which are thick walled and long-lived and formed by the Phylum Oomycota
Ootheca Egg case
Oozes Viscid masses composed of living or dead pathogen structures and partially disintegrated suscept tissues.
Operculum Lid-like structure on the eggshell
Opisthognathous Mouthparts directed posteroventrally
Opisthosoma ‘back body’, the abdomen-like region of chelicerates
Organic herbicides Herbicides made from natural materials such as salts of fatty acids and corn byproducts
Orthoptera ‘Order of isects: grasshoppers, locusts and crickets
Oviparity Development where egg is fertilised in the body and then laid.
Ovoviviparity Retention of the developing fertilised egg in the mother but there is no nutrition of the hatched young.

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Palaeoptera Insects with wings not folded over the body at rest e.g. Odonata.,
Palar pegs Found on tibia of some water insects, used to grasp female during mating.
Palpifer A sclerite on the stipes which bears the palp
Panoistic ovary Ovarioles lack trophocytes
Papillae Small, rounded processes
Papillate Bearing papillae
Paragynous Having the antheridium alongside the oogonium (typical of some Pythiaceae)
Paramere Accesory clasping structure
Paraphysis A sterile hypha in a hymenium (pl. paraphyses)
Paraproct Ventrolateral plates of the 11th abdominal segment
Pars intercerebralis Anterior mediodorsal part of the protocerebral lobes
Parthenogenesis Reproduction without fertilisation by male sperm
Pathogen Disease-causing organism
Paraglossae Outer lobes on prementum (part of labium)
Paranotal theory The hypothesis that insect wings evolved from thoracic lobes
Parapatric speciation oara = Greek for 'beside'; patria = Greek for 'country' (fatherland). Speciation between adjacent populations
Paraphyletic group An unnatural taxon in which some descendants of a common ancestor have not been included (e.g. Reptilia without Aves)
Pedial tympanum Hearing organ on the legs of Orthoptera
Pedipalps The second pair of mouthparts in chelicerates, typically palp-like, but sometimes chelate (e.g. In scorpions)
Perennial plant A plant that lives for several years
Pericardial sinus Dorsal body compartment surrounding the hear
Peridium The outer wall of a fruiting body
Peripheral nervous system Nerves that connect with the central and visceral nervous systems.
Perithecium Ascocarp which is flask-shaped with a pore (ostiole) through which ascospores escape
Peritrophic membrane Multi-layered midgut membrane which serves many functions
Permanent pastures Pastures not used in a cropping rotation, but which may be frequently or infrequently re-sown
Phallobase Proximal part of the phallus
Phallotreme Apical opening of the endophallus
Phallus Intromittent organ of the male
Pharate Concealed stage enclosed in the old cuticle of the preceding stage
Pharyngeal pump The sucking pump of some fluid feeding insects
Phasmatodea Order of insects: stick insects. Greek, phasma" = apparition, ghost
Phellogen Cork cambium
-phore (suff.) - a stalk
Photosystem I One of three major photosynthetic reaction centres. PSI creates the materials used in later photosynthetic reactions such as food production for the plant.
Photosystem II Is used in conjuction with Photosystem I. It splits water into oxygen and hydrogen atoms and is responsible for the production of atmospheric oxygen, essential for aerobic life on this planet
Phototaxis Response to light
Phragma An enlarged apodemal projection (antecosta) for muscle attachment.
Phthiraptera Order of insects: lice. Greek, phthir = lice, aptera = wingless
Phylogeny The history of life; evolutionary history of a group
Physogastric Abdomen swollen usually with eggs or larvae e.g. Termite queen
Phytohormones Plant hormones
Phytophagous Insects that feed on plants
Phytoplasma Prokaryotic organism (mollicutes) that is round to elongate in shape
Pili A structure that can aid the attachment of bacterial cells to plant surfaces; believed to be a means for genetic exchange between bacterial cells by conjugation
Pilifer Small lobe-like, often bristled remnant of the labrum located on each side at the lower angle of the face of adult Lepidoptera; it is sometimes long and touching either the proboscis or the inner face of the labial palpus.
Pit Conical invagination of the integument
Pitting Superficial depressions in the surface of succulent organs resulting in the death and shrinkage of subepidermal cells.
Placode sensillum Plate-like cuticular sense organ
Planidium Mobile first instar of some paratisitc Hymenoptera
Plasmodium (of Myxomycota) a multinucleate, naked, amoeboid body (pl. plasmodia)
Plastron Hairs or cuticular structures which hold a thin layer of gas permanently in position; functions as a physical gill
Plecoptera Order of insects: stoneflies. Greek, plekein = to fold, pteron = wing
Pleomorphic cells Cells with many shapes
Plesiomorphic Primitive condition
Pleural suture Suture dividing the pleuron into an episternum and epimeron
Pleural wing process Wing fulcrum found on the dorsal margin of the pleuron, on which the base of the wing rests
Pleurites Sclerites in the pleural region of the body wall
Pleurosternal suture Boundary between the pleuron and sternum.
Pleuron Lateral walls of the thorax
Plicae Longitudinal folds which extend the length of the proventriculus
Plumose gills Feather-like gills
Polyembryony Many individuals from one egg
Polygenic inheritance Inherited characteristics are controlled by genes at many loci located on several chromosomes. The different loci all combine to produce the final phenotype.
Polygenic resistance Resistance under the control of several genes. It is a partial resistance which is more difficult to select as it is influenced by climatic conditions
Polypneustic respiratory system Respiratory system characterised by 8 or more pairs of spiracles, and is subdivided into holopneustic, peripneustic and hemipneustic.
Polymorphism Several anatomical forms or colour variants displayed in one species
Polyphaga Suborder of Coleoptera (beetles) e.g. blister beetles, click beetles, leaf beetles
Polyphyletic The worst sin in phylogenetic systematics, an unnatural taxonomic group whose members do not share a recent common ancestor
Polypodous Larval type with 3 thoracic legs and abdominal prolegs
Polytrophic ovary Ovariole consists of alternating layers of oocytes and trophocytes
Population Plants/animals of a particular type or group who live in an area together
Pore canals Canals that extend from the epidermis to the cuticle
Postalar bridge Extension of the postnotum connecting with the epimeron
Postcoxal bridge Extension of the epimeron connecting the pleuron and sternum behind the coxa.
Postcubitus First anal vein; associated with the cubital vein
Postmentum Part of the labium attched to the head; can be divided into mentum and submentum
Posterior notal wing process Posterior articulation point for the wing
Postgena Region immediately posterior to the gena
Postgenal bridge Formed by the fusion of the postgena
Postnotum Posterior region of the notum, bearing the phragma
Postoccipital suture Suture which separates the occiput and postocciput
Postocciput Sclerite found behind the postoccipital suture; surrounds the occipital foramen
Postscutellum Rounded ridge under the mesoscutellum in some flies (Tachinidae)
Prealar bridge Externsion of the prescutum connecting with the episternum
Precoxal bridge Extension of the episternum, connects the pleuron and sternum anterior to coxa
Prementum Apical part of postmentuml bears glossae, paraglossae and palps
Prescutal suture Separates the prescutum and scutum
Prescutum Anterior sclerite of the alinotum
Presternum Small anterior sclerite of the eusternum
Pretarsus Distal-most portion of the leg
Primary pigment cell Surrounds cone and proximal portion of the lens. Pigment does not migrate
Procuticle Endocuticle and exocuticle together
Prognathous Mouthparts directed anteriorly
Prolegs Abdominal legs found in Lepidoptera and Hymenoptera
Prolepsis Premature development of a shoot from a bud.
Proliferation (prolification) Continued or extended development of a plant part after reaching a stage at which normal growth ceases.
Promycelium (of Uredinales and Ustilaginales) a short hypha which grows from a teliospore and produces basidiospores (the basidium of these orders)
Proneustic respiratory system A respiratory system characterised by a mesothoracic spiracle only e.g Diptera larvae
Pronotum Anterior dorsal sclerite of the thorax
Prophylactic Tending to prevent disease; a substance so used
Prophylactic weed control This approach aims to prevent reinfestations or symptoms from re-occurring which requires population to drastically reduced
Propodeum First abdominal segment of hymenoptera; associated with the thorax
Prosoma ‘front body’, the anterior body tagma in chelicerates, sometimes called the cephalothorax
Prothorax First segment of the thorax
Protocephalon Primitive groundplan head
Protocerebral bridge Connects the two hemispheres of the protocerebrum
Protocerebral lobes Part of the protocerebrum
Protocerebrum Anteriormost part of brain
Protura Class of entognathous hexapods
Proventriculus Part of the foregut, posterior to the crop which grinds food
Proximal (= basal) Structure or part of a structure closest to the body.
Proximal median plate Between the second and third axillary sclerites. Attaches to the medial and cubital veins.
Pseudergates Worker-like caste but not a true worker as nymphs moult into soldiers, primary reproductives or secondary reproductives
Pseudothecium Not a true ascocarp, rather the asci are formed directly in cavities within a matrix of mycelium
Pseudotracheae Ridged grooves on the ventral surface of the labellum of some flies, used to take up liquid food
Psocoptera Order of insects: booklice.
Pteral tympanum Hearing organ on the wings of Lepidoptera and Neuroptera
Pterines Pigments which produce white, yellow, red and fluorescence
Pterostigma Pigmented or defined opaque area near the distal end of the anterior margin of the wing.
Pterothorax The meso- and metathorax of winged insects, collectively
Pterygotes Winged insects
Ptilinum A sac-like, inflatable, pulsatile, cuticular organ on the head of Diptera (flies). Aids emergence from the puparium.
Pulvillus Lateral lobes of the pretarsus found beneath the claws.
Pupa Stage between larva and adult in holometabolous insects
Puparium Barrel-shaped capsule formed by hardened cuticle of final instar larva e.g. Blowflies
Pycnidia Enclosed structure with conidia on short conidiophores
Pygopods Eversible locomotory appendages on 10th abdominal segment of pterygote larvae
Pyloric valve Forms a boundary between the midgut and hindgut.

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Quarantine The process whereby the movement of plants, into and out of a country, is regulated
Quiescence Immediate response of an insect to unfavourable conditions

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Radial cross vein Connects the first branch of the radius with the anteriormost second branch of the radial sector
Radial vein Third vein; articulates with the second axillary sclerite
Raptorial Legs adapted for prey capture
Receptive hypha One of the sex organs of Uredinales - accepts spermatia
Reddening The loss of normally developed green colour from chlorophyllous tissue due to the destruction of the chlorophyll and/or degeneration of the chloroplasts which leads to a display of red pigments.
Remigium Anterior surface area of the wing; envelops most of the large wing veins, relatively rigid area.
Reniform Bean-shaped
Repagula Rod-like bodies expelled from the female during oviposition
Resilin Rubber-like or elastic protein found in some insect cuticle
Resinosis Flow of resin or pitch from wounds or other lesions in conifers.
Restoration Development of tissues or organs that normally would be only rudimentary.
Retinaculum A hook-like holdfast structure found on the 3rd abdominal segment of Collembola; part of the "jumping" organ
Retinular cell Sensory cell of the ommatidium
Rhabdomere Formed on the margin of each retinular cell and contains the visual pigment. All the rhabdomeres together form the rod-like rhabdom
Rhizoids Branched hyphal filaments of limited length which anchor some fungi and are also used for the absorption of nutrients and water
Rhizosphere The region in the vicinity of the root
Roguing Removal of diseased plants or plants of a variety different from that planted in the field
Rosetting A shortening of internodes through failure to elongate, which results in a crowding of the foliage.
Rot (decay) Disintegration and decomposition of dead tissue.
Russeting Rough or corky surfaces formed where they do not normally occur.
Rust Important group of the plant parasites. The name comes from the fact that the sori and spores, are yellow-orange in colour

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Sachet Fluid excretion encapsulated within a membrane-like sac
Sacculus Pit of the coeloconic sensillum
Safener A chemical added to a pesticide to keep it from injuring plants>
Salivarium Ventral area of preoral cavity (formed by the hypopharynx)
Saltatorial Adapted for jumping
Saprogenesis The period in the life cycle of a pathogen when it is not associated with living tissues of its host i.e. it is sapophytic. This may include a period of dormancy
Saprophagy ‘putrid eating’, feeding on dead things
Saprophyte A plant living on dead or decaying organic matter
Sarcody Swelling of bark beyond girdling wounds or cankers. .
Scab A limited, more or less circular, raised and sometimes roughened lesion on fruits, tubers, leaves and stems, resulting from an overgrowth of epidermal, cortical, and peridermal tissues.
Scaling Formation of scales or corky material where scales normally don't occur.
Scansorial Adapted for climbing
Scarabaeiform Larval type which is C or U-shaped
Sclerites Hardened areas of the insect body wall also called "plates"
Sclerotes Hard resting structures containing toxins, produced by the fungus which allow it to survive adverse conditions
Sclerotium A firm, frequently rounded, mass of hyphae with or without incorporated host tissue or soil, and having no spores in or on it (pl. sclerotia)
Sclerotin Structural protein that forms the hard part of the integument
Scolophorous organs Sense organs with an auditory function
Scorch A sudden drying and browning of large, indefinite areas on leaves and fruits. Also damage to bark resulting in drying and death.
Scototaxis Response to darkness
Scutoscutellar suture Suture which separates the scutum and the scutellum
Scutum Mid sclerite of the alinotum
Scutellum Posterior sclerite of the alinotum
Sectorial cross vein Connects two medial branches of radial sector
Second axillary sclerite Pivotal plate of wing base, rests on the pleural wing process, connected with the radial vein
Secondary pigment cell Separates ommatidia. Pigment can migrate or it may be restricted in the cell.
Seminal vesicle Enlargement of the vas deferens which stores sperm
Senescent Declining or dying
Sensilla Simplest sense cells formed from epidermal cells
Sensillum Sense organ found on tergum 10 of fleas
Sensory neurones Associated with sensory structures
Septate hypha Hyphae which have septa
Septum Cross wall of a hypha, spore etc (pl. septa) that has a central pore to allow cytoplasmic continuity throughout
Serology Method for identifying microorganisms based on the fact that an organism may be used as an antigen
Shelling Large scale loss of leaves through premature abscission.
Shot-hole Holes in leaves caused by the prompt falling out of the killed spots of tissue.
Silvering A silvery sheen or gray luster of leaf or primary bark surfaces due to the development of unnatural air spaces beneath the epidermis.
Siphonaptera Order of insects: fleas.
Somites Body segments
Sorus A spore mass (especially Uredinales and Ustilaginales) (pl. sori)
Speciation The origination of new species
Spermatium A + or - sex cell (especially of Uredinales) (pl. spermatia)
Spermatogenesis Formation of spermatozoa
Spermatagonia Male germ cells that divide and form spermatocytes
Spermatazoa Cells that divide and become the spermatazoa
Spermatheca A sac which stores sperm inside the female
Spermatids Developing male cells formed by division of spermatocytes
Spermatocytes Cells that divide and become the spermatazoa
Spermatophore Container which protects, envelopes and transfers sperm from the male to the female
Sphragis Mating plug in Lepidoptera
Spicisetae Setae composed of sharply pointed overlapping scales
Spike-top Death of the entire crown of the tree in a pattern that resembles staghead.
Spinasternum Smaller, posterior plate of sternum
Spiracles A pore in the integument which serves for gas exchange
Spiroplasma Prokaryotic organism (mollicutes) that is helical (spiral) in shape
Sporangium A container that holds asexual spores (pl. sporangia) usually found in the Lower Fungi
Spore General term for the reproductive units of fungi that consist of one or a few cells
Sporidium A basidiospore of the uredinales or ustilaginales (pl. sporidia)
Spot Circular, areolate, or irregular discoloured and dead areas on leaves, fruits or green stems..
Spray-topping Application of low rates of herbicide to weeds prior to or in early flowering to substantially reduce the number of viable seeds set by plants and hence the soil seed bank for future years
Spray-grazing The application of sub-lethal doses of herbicides e.g. glyphosate, to weeds to increase their sugar content and palatability, thereby encouraging livestock to graze them in preference to other pasture plants
Spumaline Froth which used t envelop insects and make it invisible
Squamae (alula, calypters). Pair of membranous lobes found in the posterobasal portion in some Diptera.
Stage Synonymous with the term instar
Stadium Interval of time between moults or the duration of an instar
Staghead Dieback or flagging of entire large branches in the crown, or of the entire top of the tree.
Stemmata Larval visual organs in the head region where the compound eye will develop
Sternites Sclerites of the ventral part of the body wall (sternum)
Sternacostal suture Transverse suture dividing the mediosternite into a basisternum and sternellum.
Sternellum Posterior sclerite of the mediosternite.
Sternorrhyncha Suborder of the Hemiptera (bugs) e.g. coccids, aphids, scale insects.
Sternum The sclerotized ventral area of the thorax
Stipes Large segment of maxilla; bears galea and lacinia
Stomodeal nervous system Part of visceral nervous system; associated with the brain, aorta and foregut.
Stomodaeal (cardiac) valve Sphincter which controls the flow of food into the midgut
Streak Elongated, narrow, superficial lesions which are first waterSoaked then brown in foliage or green stems. In wood there is discolouration in axial streaks within the outer, conducting layer of sapwood resulting from deterioration of the wood structure or extraneous components.
Strepsiptera Order of insects. Greek, streptos = twisted, pteron = wing
Stridulation Production of sound by friction
Stripe Streak symptom in monocots.
Stunting A type of dwarfing in which the entire plant is subnormal in size.
Stylets Elongate, narrow modifications of generalised, chewing mouthparts
Styloconic sensillum Tooth or peg-like sense organ
Stylus Small abdominal appendage
Subalare A sclerite at the base of the wing, posterior of the pleural wing process
Subcostal vein Second vein; articulates with the first axillary sclerite
Subgena Narrow area above the mouthparts.
Subimago Stage between the naiad and adult. Subimago has functional wings and undergoes ecdysis to produce the adult stage
Suboesophageal ganglion Anteriormost ganglion of the ventral nerve cord
Succession The change of plant communties over time
Sulcus Externally visible line formed by the inflection of cuticle
Supernumerary sperm Sperm not used in fertilisation
Superposition eye Forms an image when light reaching one rhabdom is transmitted from adjacent ommatidia. Found in nocturnal, crepuscular insects
Suppression Complete prevention of organ development.
Sutures Seams produced by the union of adjacent sclerites
Symbiont An organism having a mutually beneficial relationship with a dissimilar organism
Symmetry Type or order where an object will coincide with its mirror image
Sympatric speciation Sym = Greek for 'together'; patria = Greek for 'country' (fatherland). Origin of a new species within the range of its ancestor
Symphyta Suborder of the Order Hymenoptera (ants, bees and wasps)
Sympleisomorphies Shared primitive characters, often useful in diagnoses, but not capable of defining monophyletic groups, e.g. Insects have antennae, but so do other hexapods, myriapods, etc.
Synapomorphies Shared derived characters, the only characters that can be used to support monophyletic taxa, e.g. Pterygote insects are the only arthropods to have wings
Systematics The study of the classification of organisms, their relationships and evolutionary history
Systemic Throughout the body

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Tabtoxin A Low molecular weight bacterial toxin produced by Pseudomonas syringae pathovars which affects normal metabolism of the host
Taenidia Cuticular rings which strengthen the trachea
Tagma Discrete units of an arthropod’s exoskeleton formed by the fusion of segments into functional units (e.g. thorax) (pl. Tagmata)
Tagmosis Organisational process where somites fuse to form distinct tagma e.g. Head, thorax, abdomen
Tarsus Most distal part of the leg which is broad and flat, divided into several "segments" (tarsomeres).
Taxonomy The science of naming and identifying organisms.
Tectiform Wings held in a roof-like position over the body when at rest for e.g. booklice (Order Psocoptera)
Tegmen Leathery forewing found in Orthoptera, Mantodea and Blattodea. Protects the hindwing when at rest
Tegula Lobe-like prothoracic structure covering the base of the forewings
Teliospore The spore of the Uredinales and Ustilaginales from which the basidium is produced; initially dikaryotic
Teletoky Production of females from unfertilised eggs (no males)
Telopodite Remainder of the leg segments
Telotrophic ovary Trophocytes are confined to the apex of each ovariole
Teneral Describes the adult shortly after emergence; integument is not hardened or fully coloured
Tentorium Internal cuticular framework of the head
Tergites Sclerites of the dorsal part of the body (tergum)
Thanatosis Feigning death
Therapeutic weed control Approach that aims to suppress an infestation or symptom, attemps to reduce losses by treating infestation in the current crop.
Thigmotactic Species that live in close proximity or in contact with a surface or in a crevice. Movement is inhibited by contact with sufaces
Third axillary sclerite The flexor sclerite of the wing base, articulates with the anal vein(s).
Thysanoptera Order of insects: thrips. Greek, thysanos = fringe, pteron = wing
Thysanura Class of entogonathous hexapods: silverfish. Greek, thysanos = fringe, oura = tail
Tibia Fourth segment of the leg ("shin")
Tillage Disturbance of soil by mechanical means usually as a preparation for sowing of crops or pastures
Tonofibrillae Cuticular fibrils connecting muscles to the integument
Tormagen cell Socket for the trichogen cell
Trachea Largest tubes of the respiratory system
Tracheoles Smaller tubules of the tracheal system
Trichoptera Order of insects: caddisflies. Greek, Thrix = hair, pteron = wing
Trichogen cell Cuticular sensory recpetive portion of the trichoid sensillum
Trichosor A setose thickening of the wing membrane along the margin
Tritocerebrum Posteriormost part of the brain
Triturating surface Grinding surface in the proventriculus
Triungulin An active, dispersive first-instar larva
Triungulinid Mobile first instar larva of Strepsiptera
Trochanter Basal segment of the telepodite and second of the leg.
Trochantin Small sclerite at the base of the leg
Trophic level The level at which energy in a food web resides, e.g. Primary producers (green plants) produce energy and therefore reside on the 1st trophic level, primary consumers (herbivores) hold energy at the 2nd trophic level
Trophocytes Nurse cells of the ovary or testis
Trophollaxis Exchange of food between insects of the same or different species
Tumefaction Local swelling on any part of the plant, usually woody roots, stem, or branches and usually resulting from stimulation of the plant meristem by the pathogen.
Turbinate eyes An eye on a cylindrical stalk
Tympanal organ "Ears" of insects
Tympanum Insect "eardrum"

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Uniramous With one branch, as in insect legs
Univoltine Having one generation in one year

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Valvifer Proximal structures of the ovipositor that bear the valvulae
Valvulae Distal processes of the ovipositor borne on the valvifers
Vannus or anal region Separated from the remigium by an anal (vannal) fold. Often triangular, best expressed in the posterior wing.
Vas deferens Duct which connects the testis to the seminal vesicle
Vector Arthropod which transmits a pathogen to plants and animals
Ventral visceral nervous system Nervous system associated with the ganglia of the ventral nerve cord
Ventriculus The true stomach of an insect
Verrucae Elevated tubercles bearing setae on caterpillars
Vertex Dorsal region of the head between the eyes
Virescence (greening) Development of chlorophyll where it is not normally present.
Visceral nervous system Innervates the gut, reproductive organs and tracheal system
Vitelline membrane Innermost layer of the eggshell
Vitellarium Region of the ovariole where vitellogenesis occurs
Vitellogenesis Deposition of egg yolk
Viviparity Eggs develop within the female; immatures laid as larvae or nymphs
Vulva Point of aedeagal intromission

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glossary icon - W

Weeds A plant existing at places and/or times in which they are considered undesirable
Weed management The plan devised for controlling the weed population so as to reduce the impact the weed has on human activity
Wilting A flaccid appearance of the leaves and shoots resulting from a temporary or permanent loss of turgor due to excess transpiration by the leaves and shoots.
Witches' broom A type of fasciculation in which there is an abnormal bushlike development of many weak shoots or roots.

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glossary icon - X

Xylophagous Wood feeding insect

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glossary icon - Y

Yellowing Loss of normally developed green colour due to the destruction of the chlorophyll and/or degeneration of the chloroplasts which leads to the display of yellow pigments.

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glossary icon - Z

Zoospores Motile spores which need water to move and be discharged
Zygoptera Suborder of Odonata (dragonflies), the damselflies.
Zygospores Sexual spores which are thick walled and formed by fungi in the Phylum Zygomycota

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